Ryan, M.J., Evans, D.C., Currie, P.J., Brown, C.M. & Brinkman, D. (2011) New leptoceratopsids from the Upper Cretaceous of Alberta, Canada. Cretaceous Research (advance online publication) doi:10.1016/j.cretres.2011.11.018

Abstract

Two new leptoceratopsid neoceratopsians are described based on partial dentaries collected from the Dinosaur Park (Campanian) and Milk River (Santonian) formations of Alberta. The new Campanian taxon has a unique dentary tooth shape not shared by other leptoceratopsid taxa, which has implications for the evolution of the Leptoceratopsidae. The Santonian specimen represents the oldest known leptoceratopsid (∼83 Ma), and probably represents the smallest adult-sized ceratopsian known from North America.


Unescoceratops koppelhusae Dinosaur Park Alberta Canada (Campanian)
Holotype: TMP 95.12.6, a partial left dentary.
Locality and horizon: Black Coulee locality (formerly Deadhorse Coulee), near the Writing-on-Stone Provincial Park, from the Dinosaur Park Formation, Alberta, Canada (Campanian)
Unescoceratops shows a unique dentary tooth morphology among other leptoceratopsids



Gryphognathus morrisoni Milk River formation Alberta Canada (Santonian)
Holotype: ROM 56635, a partial right dentary
Locality and Horizon: Dinosaur Provincial Park, fbonebed 55, Milk River Formation,late Santonian, Alberta Canada