[N] 2004 First direct fossil evidence that tyrannosauroids had protofeathers
20041012
D.paradoxus is the earliest known unquestionable tyrannosauroid found so far (2004). One of the specimens also preserves a filamentous integumentary covering similar to that of other coelurosaurian theropods from western Liaoning. This provides the first direct fossil evidence that tyrannosauroids had protofeathers