Colorado - US

The Laramie formation Colorado, US was deposited by low-lying coastal swamps and estuaries along the edge of the Cretaceous sea. Coal seams are common in the lower part of the formation. Typical fossils include palm leaves and dinosaur bones. The Laramie Formation contains multiple track-bearing horizons exhibiting a diverse assemblage of ceratopsid, ornithopod and theropod tracks and trackways.

The ceratopsid tracks are the first attributable to this family and we describe them as a new ichnotaxon, Ceratopsipes goldenensis, ichnogen. et ichnosp. nov. The ceratopsid-dominated ichnofauna reflects the body fossil record of the Laramie Formation.

Theropod tracks indicate a diverse fauna including an ichnite that possibly represents Tyrannosaurus. A partial trackway represents the first reported champsosaur ichnites herein named Champsosaurichnus parfeti ichnogen. et ichnosp. nov. The ichnofauna also includes possibly hadrosaur tracks.


Agathaumas milo - Edmontosaurini Nomen Nudem - Cretaceous Late Maastrichtian


Claosaurus annectens - Hadrosaurinae Edmontosaurini - Cretaceous Late Maastrichtian



Dipriodon caperatus - Troodontidae Nomina Dubia - Cretaceous Late Maastrichtian



Edmontonia sp (1) - Not considered dinosaur



Edmontosaurus regalis - Hadrosaurinae Edmontosaurini - Cretaceous Late Maastrichtian



\\\"Meniscoessus\\\" caperatus - Troodontidae Nomina Dubia - Cretaceous Late Maastrichtian



Ornithomimus minutus ~/~ - Avialae Nomina Dubia - Cretaceous Late Maastrichtian



Paronychodon caperatus ? - Troodontidae Nomina Dubia - Cretaceous Late Maastrichtian



Thescelosaurus neglectus - Ornithopoda Hypsilophodontidae - Cretaceous Late Campanian Maastrichtian



Torosaurus latus - Ceratopidae Chasmosaurinae - Cretaceous Late Maastrichtian



Triceratops horridus - Ceratopidae Chasmosaurinae - Cretaceous Late Maastrichtian



Tripriodon caperatus - Troodontidae Nomina Dubia - Cretaceous Late Maastrichtian



Triprotodon carperatus - Troodontidae Nomina Dubia - Cretaceous Late Maastrichtian



Ceratopian footprints - Ceratopia - Cretaceous Late Maastrichtian



Ornithopod footprints - Ornithopoda - Cretaceous Late Maastrichtian



Theropod footprints - Theropoda - Cretaceous Late Maastrichtian



?Dromaeosaurid indet. - Theropoda - Cretaceous Late Maastrichtian



?Edmontonia sp - Ankylosauria Nodosauridae - Cretaceous Late Maastrichtian



cf. Tyrannosaurus rex - Theropoda Tyrannosauridae - Cretaceous Late Maastrichtian



Thescelosaurus sp. - Ornithopoda Hypsilophodontidae - Cretaceous Late Maastrichtian



Triceratops sp. - Ceratopia Ceratopidae - Cretaceous Late Maastrichtian



Troodontid indet. (= Paronychodon sp.) - Theropoda - Cretaceous Late Maastrichtian