Galve, Teruel Province, Spain

The Villar del Arzobispo Formation is a regressive 100 m-thick succession of marine limestones, sandstones and unconsolidated mudstones, with an association of foraminiferans including Pseudocyclamina gr. lituus, Archispirocyclina lusitanica and Rectocyclamina arrabidensis which give a Tithonian age at the top of this unit (Díaz and Yébenes, 1987). The formation spans the late Tithonian and the early and mid Berriasian. Two fossil sites are known towards the top of this unit: Cuesta Lonsal, source of the remains of a new cetiosaurid sauropod dinosaur (Sánchez-Hernández, 2005) and associated teeth and scales of fishes; and Las Cerradicas, which yielded the footprints of theropods and ornithopods, including the trail of an iguanodontian dinosaur that started with a quadrupedal gait and finished as a biped (Pérez-Lorente et al., 1997; this outcrop was initially dated as Berriasian, but Cuenca-Bescós et al. (1997) revised its age to Late Jurassic). There are also many ripples in these outcrops generated under fluvial influence. In similar beds, or slightly lower, is the El Cantalar outcrop, which produced footprints of an enormous crocodyliform with a total length of about 12 m, although its taxonomic attribution is unclear (Pérez-Lorente and Ortega, 2003). [Sánchez-Hernández et al. 2007]


Galveosaurus herreroi - Sauropodomorpha Sauropoda - Jurassic Late Cretaceous Early Tithonian Berriasian


Turiasaurus riodevensis - Sauropoda Turiasauria - Jurassic Late Cretaceous Early Tithonian Berriasian