Churcher, C. S., and de Iuliis, G. (2001) A new species of Protoperus and a revision of Ceratodus humei (Dipnoi: Ceratodontiformes) from the Late Cretaceous Mut Formation of Eastern Dakhleh Oasis, western Desert of Egypt: Palaeontology, v. 44, part 2, p. 305-323. (Protopterus crassidens)